Abstract:The continuous Permian-Triassic strata in West Qinling represent deep-marine deposits. In this paper,abundant trace fossils(seventeen ichnospecies,twelve ichnogenera)have been identified in the Upper Permian Maomaolong Formation in the Gannan area of the western Qinling orogen. As a new ichnologic method,ichnonetwork analysis focuses on the reconstruction of the community system composed of interconnected ichnotaxa,which provides useful means for studying the behavior and ecology of ichnocoenosis. In this study,an ichnonetwork analysis is conducted with trace fossils data obtained from the first hand filed trip. The results show that ichnocoenosisis structured in the Maomaolong Formaiton,and three ichnoassociations were controlled by palaeoenvironments: Palaeophycus-Lockeia-Arenicolites-Dictyodora-Planolites-Protovirgularia-Chondrites (C1,proximal turbidite zone),Palaeophycus-Lockeia-Arenicolites-Dictyodora-Protovirgularia-Phycodes-Chondrites-Helminthopsis (C2,turbidite transition zone-distal turbidite zone),and Palaeophycus-Nereites-Scolicia-Protovirgularia-Helminthopsis (C3,distal turbidite zone). The evolution from C1 to C3 in the Upper Permian Maomaolong Formation from the Gannan area of the West Qinling reveals the palaeoenvironmental changes from proximal turbidite zone to distal turbidite zone.
Dang Zhi-Ying,Zhang Li-Jun. Palaeoenvironmental evolution of the Upper Permian Maomaolong Formation in Gannan area of the West Qinling: Based on ichnonetwork analysis[J]. JOPC, 2020, 22(6): 1171-1180.
[1] 方国庆,张晓宝,李育慈. 1994. 西秦岭华力西—印支期幕式裂陷活动初探. 西北地质, 15(1): 1-5. [Fang G Q,Zhang X B,Li Y C.1994. Rifi-fauitings of Hercynian-Indosinan epoch of western Qingling orogenic belt and its relationship with Palaeotethys. Northwestern Geology, 15(1): 1-5] [2] 寇晓虎,朱云海,张克信,陈中强,施彬,罗根明. 2007. 青海省同仁地区上二叠统石关组上部火山岩的新发现及其地球化学特征和构造环境意义. 地球科学, 32(1): 45-50,58. [Kou X H,Zhu Y H,Zhang K X,Chen Z Q,Shi B,Luo G M.2007. Discovery and geochemistry of Upper Permian volcanic rocks in Tongren area,Qinghai Province and their tectonic significance. Earth Science, 32(1): 45-50,58] [3] 李柒林,代文军,刘生祥,邱庆伦. 2002. 甘肃省夏河地区二叠系毛毛隆组砾屑灰岩成因讨论. 沉积与特提斯地质, 22(3): 85-89. [Li Q L,Dai W J,Liu S X,Qiu Q L.2002. Discussion on genesis of conglomerate limestone of Permian Mao Maolong Formation in Xiahe area,Gansu Province. Sedimentary Geology and Tethyan Geology, 22(3): 85-89] [4] 罗根明,张克信,林启祥,寇晓虎,朱云海,徐亚东,施彬. 2007. 西秦岭地区晚二叠世—早三叠世沉积相分析和沉积古环境再造. 沉积学报, 25(3): 332-342. [Luo G M,Zhang K X,Lin Q X,Kou X H,Zhu Y H,Xu Y D,Shi B.2007. Sedimentary facies analysis and sedimentary environment reconstruction from Late Permian to Early Triassic of West Qinling area. Acta Sedimentologica Sinica, 25(3): 332-342] [5] 王鸿祯. 1985. 中国古地理图集. 北京: 地图出版社. [Wang H Z. 1985. Atlas of Chinese Paleogeography. Beijing: Cartographic Publishing House] [6] 杨式溥,张建平,杨美芳. 2004. 中国遗迹化石. 北京: 科学出版社,1-353. [Yang S P,Zhang J P,Yang M F.2004. Trace Fossils of China. Beijing: Science Press,1-353] [7] 殷鸿福,杨逢清,黄其胜,赖旭龙. 1992. 秦岭及邻区三叠系. 武汉: 中国地质大学出版社. [Yin H F,Yang F Q,Huang Q S,Lai X L. 1992. The Triassic of Qinling Mountains and nieghboring areas. Wuhan: China University of Geosciences Press] [8] 赵江天,杨逢清. 1992. 甘肃夏河麻隆沟二叠纪毛毛阻组沉积特征及环境分析. 沉积学报, 10(1): 47-56. [Zhao J T,Yang F Q.1992. Permian sedimentary features and environmental analysis of Maomaolong Formation in Malonggou Xiahe,Gansu. Acta Sedimentologica Sinica, 10(1): 47-56] [9] 左国朝,金松桥,朱伟元. 1984. 甘肃夏河县下卡加—完孕滩一带二叠系浊积岩及有关粗碎屑沉积物. 沉积学报, 2(3): 75-82. [Zuo G Z,Jin S Q,Zhu W Y.1984. Permian turbidites and related coarse clastic sediments in Xiakajia-Wangatan area of Xiahe County,Gansu Province. Acta Sedimentologica Sinica, 2(3): 75-82] [10] Barabsi A L.2002. Linked: The New Science of Networks. New York: Perseus Books Group,280. [11] Bascompte J.2007. Networks in ecology. Basic and Applied Ecology, 8: 485-490. [12] Baucon A,Felletti F.2013a. Neoichnology of a barrier-island system: The Mula di Muggia(Grado lagoon,Italy). Palaeogeography,Palaeoclimatology,Palaeoecology, 375: 112-124. [13] Baucon A,Felletti F.2013b. The IchnoGIS method: Network science and geostatistics in ichnology. Theory and application(Grado lagoon,Italy). Palaeogeography,Palaeoclimatology,Palaeoecology, 375: 83-111. [14] Baucon A,Ronchi A,Felletti F,de Carvalho C N.2014. Evolution of Crustaceans at the edge of the end-Permian crisis: Ichnonetwork analysis of the fluvial succession of Nurra(Permian-Triassic,Sardinia,Italy). Palaeogeography,Palaeoclimatology,Palaeoecology, 410: 74-103. [15] Baucon A,Venturini C,de Carvalho C N,Felletti F,Muttoni G.2015. Behaviors mapped by new geographies: Ichnonetwork analysis of the Val Dolce Formation(lower Permian;Italy-Austria). Geosphere, 11(3): 744-776. [16] Benton M J,Trewin N J.1980. Dictyodora from the Silurian of Peeblesshire,Scotland. Palaeontology, 23(2): 501-513. [17] Bradley J.1980. Scolicia & Phycodes,trace fossils of Renill(Pennatulacea). Pacific Geology, 14: 73-86. [18] Bradley J.1981. Radionerites,Chondrites and Phycodes,trace fossils of Athoptiloid sea pens. Pacific Geology, 15: 1-16. [19] Bromley R G.1996. Trace fossils: Biology,taphonomy and applications. Palaeogeography,Palaeoclimatology,Palaeoecology, 129(1): 193-194. [20] Buatois L A,Mngano M G.2011. Ichnology: Organism-substrate interactions in space and time. New York,Cambridge:Cambridge University Press. [21] Carmona N B,Mngano M G,Buatois L A,Ponce J J.2010. Taphonomy and paleoecology of the bivalve trace fossil protovirgularia in deltaic heterolithic facies of the Miocene Chenque Formation,Palagonla,Argentina. Journal of Paleontology, 84(4): 730-738. [22] Crimes T P.1987. Trace fossils and correlation of Late Precambrian and Early Cambrian strata. Geological Magazine, 124(2): 97-119. [23] Ekdale A A,Bromley R G.1984. Comparative ichnology of shelf-sea and deep-sea chalk. Journal of Paleontology, 58(2): 322-332. [24] Ekdale A A,Mason T R.1988. Characteristic trace-fossil associations in oxygen-poor sedimentary environments. Geology, 16(8): 720-723. [25] Feng X Q,Chen Z Q,Michael J B,Wu S Q,David J B,Jeffrey R T.2019. A diverse trackway-dominated marine ichnoassemblage from the Lower Triassic in the northern Paleotethys: Ichnology and implications for biotic recovery. Palaeogeography,Palaeoclimatology,Palaeoecology, 519: 124-140. [26] Fillion D,Pickerill K R,Harland T L.1990. Influence de la diagenèse sur la production,la préservation et l,échantillonnage des ichnofossiles au sein des séries carbonatées: L,exemple du bassin intracratonique ordovicien du Lac-St-Jean et de Chicoutimi(Québec). Geobios, 23(4): 485-511. [27] Gingras M K,MacEachern J A,Dashtgard S E.2011. Process ichnology and the elucidation of physico-chemical stress. Sedimentary Geology, 237(3-4): 115-134. [28] Hammer O,Harper D.2006. Paleontological Data Analysis. Blackwell,Malden. [29] Häntzschel W.1975. Trace fossils and problematica(2nd edition). In: Teichert C(ed). Treatise on Invertebrate Paleontology,Miscellanea,Supp 1. Lawrence. Geological Society of America. [30] Jaccard P.1901. Etude comparative de la distribution florale dans une portion des Alpes et des Jura. Bulletin de la Societe Vaudoise des Sciences Naturelles, 37(142): 547-579. [31] Jaccard P.1912. The distribution of the flora in the alpine zone. New Phytologist, 11(2): 37-50. [32] Kou X H,Zhang K X,Zhu Y H,Chen Z Q,Lin,Q X,Chen F N,Huang J Y.2009. Middle Permian seamount from Xiahe area,Gansu Province,northwest China: Zircon U-Pb age,biostratigraphy and tectonic implications. Journal of Earth Science, 20(2): 364-380. [33] Lewis T G.2009. Network Science: Theory and Applications. Hoboken,New Jersey:John Wiley & Sons,524. [34] Mngano M G,Buatois L A,West R R,Maples C G.1998. Contrasting behavioral and feeding strategies recorded by tidal-flat bivalve trace fossils from the Upper Carboniferous of eastern Kansas. Palaios, 13(4): 335-351. [35] Montoya J M,Pimm S L,Solé R V.2006. Ecological networks and their fragility. Nature, 442: 259-264. [36] Narbbonne G M.1984. Trace fossils in Upper Silurian tidal flat to basin slope carbonate of Arctic Canada. Journal of Paleoentology, 58(2): 398-415. [37] Osgood R G.1970. Trace fossils of the Cincinnati area. Paleontographica Americana, 6(41): 281-444. [38] Pemberton S G,Spila M,Pulham A J,Saunders T,MacEachern J A,Robbins D,Sinclair I K.2001. Ichnology and sedimentology of shallow to marginal marine systems: Ben nevis and avalon reservoirs,jeanne d,Arc Basin. Geological Association of Canada: Short Course Notes. [39] Pemberton S G,Frey R W.1982. Trace fossil nomenclature and the Planolites-Palaeophycus dilemma. Journal of Paleontology, 56(4): 843-881. [40] Reineck H E.1967. Parameter von Schichtung und Bio_turbation. Geologische Rundschau, 56(1): 420-438. [41] Seilacher A.1964. Biogenic Sedimentary Structures. In: Imbrie J,Newell N(eds). Approaches to Paleoecology. New York: Wiley,296-316. [42] Seilacher A.1994. How valid is Cruziana stratigraphy?International Journal of Earth Sciences, 83(4): 752-758. [43] Shi G R.2006. The marine Permian of East and Northeast Asia: An overview of biostratigraphy,palaeobiogeography and palaeogeographical implications. Journal of Asian Earth Sciences, 26(3-4): 175-206. [44] Smith A B,Crimes T P.1983. Trace fossils formed by heart urchins a study of Scolicia and related traces. Lethaia, 16(1): 79-92. [45] Uchman A.1998. Taxonomy and ethology of flysch trace fossils: Revision of the Marian Ksiazkiewicz collection and studies of complementary materia. Annales Societatis Geologorum Poloniae, 68(2-3): 105-218. [46] Uchman A.2004. Phanerozoic history of deep-sea trace fossils. Geological Society,London,Special Publications, 228(1): 125-139. [47] Wassermann S,Faust K.1994. Social Network Analysis. Cambridge: Cambridge University Press. [48] Wetzel A,Uchman A.1997. Ichnology of deep-sea fan overbank deposits of the Ganei slates(Eocene,Switzerland): A classical flysch trace fossil locality studied first by Oswald Heer. Ichnos, 5(2): 139-162. [49] Wetzel A,Uchman A.2012. Hemipelagic and pelagic basin plains. In: Knaust D,Bromley R G(eds). Trace fossils as indicators of sedimentary environments. Developments in Sedimentology 64. Elsevier Amsterdam. [50] Wetzel A.2002. Modern Nereites in the South China Sea: Ecological association with redox conditions in the sediment. Palaios, 17(5): 507-515.