a New Mexico Museum of Natural History, 1801 Mountain Road NW, Albuquerque, New Mexico 87104-1375 USA; b Department of Biological Sciences, Le Moyne College, 1419 Salt Springs Road, Syracuse, NY 13214 USA
The Late Triassic was a prolonged interval of elevated extinction rates and low origination rates that manifested themselves in a series of extinctions during Carnian, Norian and Rhaetian time. Most of these extinctions took place in the marine realm, particularly affecting radiolarians, conodonts, bivalves, ammonoids and reef-building organisms. On land, the case for a Late Triassic mass extinction is much more tenuous and has largely focused on tetrapod vertebrates (amphibians and reptiles), though some workers advocate a sudden end-Triassic (TJB) extinction of land plants. Nevertheless, an extensive literature does not identify a major extinction of land plants at the TJB, and a comprehensive review of palynological records concluded that TJB vegetation changes were non-uniform (different changes in different places), not synchronous and not indicative of a mass extinction of land plants. Claims of a substantial perturbation of plant ecology and diversity at the TJB in East Greenland are indicative of a local change in the paleoflora largely driven by lithofacies changes resulting in changing taphonomic filters. Plant extinctions at the TJB were palaeogeographically localized events, not global in extent. With new and more detailed stratigraphic data, the perceived TJB tetrapod extinction is mostly an artifact of coarse temporal resolution, the compiled correlation effect. The amphibian, archosaur and synapsid extinctions of the Late Triassic are not concentrated at the TJB, but instead occur stepwise, beginning in the Norian and extending into the Hettangian. There was a disruption of the terrestrial ecosystem across the TJB, but it was more modest than generally claimed. The ecological severity of the end-Triassic nonmarine biotic events are relatively low on the global scale. Biotic turnover at the end of the Triassic was likely driven by the CAMP (Central Atlantic Magmatic Province) eruptions, which caused significant environmental perturbations (cooling, warming, acidification) through outgassing, but the effects on the nonmarine biota appear to have been localized, transient and not catastrophic. Long-term changes in the terrestrial biota across the TJB are complex, diachronous and likely climate driven evolutionary changes in the context of fluctuating background extinction rates, not a single, sudden or mass extinction.
.Abdala, F., Ribeiro, A. M., 2010. Distribution and diversity patterns of Triassic cynodonts (Therapsida, Cynodontia) in Gondwana. Palaeogeography, Palaeoclimatology, Palaeoecology, 286, 202-217.
[2]
.Ash, S., 1986. Fossil plants and the Triassic-Jurassic boundary, in: Padian, K. (Ed.), The Beginning of the Age of Dinosaurs. Cambridge University Press, Cambridge, pp. 21-30.
[3]
.Avanzini, M., Pinuela, L., Garcia-Ramos, J. C., 2010. First report of a Late Jurassic lizard-like footprint (Asturias, Spain). Journal of Iberian Geology, 36, 175-180.
[4]
.Bacon, K. L., Belcher, C. M., Haworth, M., McElwain, J. C., 2013. Increased atmospheric SO2 detected from changes in leaf physiognomy across the Triassic-Jurassic boundary interval of East Greenland. PLOS One, 8(4), e60614.
[5]
.von Baczko, M. B., Ezcurra, M. D., 2013. Ornithosuchidae: A group of Triassic archosaurs with a unique ankle joint, in: Nesbitt, S. J., Desojo, J. B., Irmis, R. B., (Eds.), Anatomy, phylogeny and palaeobiology of Early Archosaurs and their kin. Geological Society, London, Special Publications, 379, 187-202.
[6]
.Baird, D., 1986. Some Upper Triassic reptiles, footprints, and an amphibian from New Jersey. The Mosasaur, 3, 125-153.
[7]
.Balini, M., Lucas, S. G., Jenks, J. M., Spielmann, J. A., 2010. Triassic ammonoid biostratigraphy: An overview, in: Lucas, S. G. (Ed.), The Triassic timescale. Geological Society, London, Special Publications, 334, 221-262.
[8]
.Bambach, R. K., Knoll, A. H., Wang, S. C., 2004. Origination, extinction, and mass depletion of marine diversity. Paleobiology, 20, 522-542.
[9]
.Bandyopadhyay, S., Sengupta, D. P., 2006. Vertebrate faunal turnover during the Triassic-Jurassic transition: An Indian scenario. New Mexico Museum of Natural History and Science Bulletin, 37, 77-85.
[10]
Belcher, C. M., Mander, L., Rein, G., Jervis, F. X., Haworth, M., Hesselbo, S. P., Glasspool, I. J., McElwain, J. C., 2010. Increased fire activity at the Triassic/Jurassic boundary in Greenland due to climate-driven floral change. Nature Geoscience, 3, 426-429.
[11]
Benton, M. J., 1986. More than one event in the Late Triassic mass extinction. Nature, 321, 857-861.
[12]
Benton, M. J., 1991. What really happened in the Late Triassic? Historical Biology, 5, 263-278.
[13]
Benton, M. J., 1993. Reptilia, in: Benton, M. J., (Ed.), The Fossil Record 2. Chapman and Hall, London, pp. 681-715.
[14]
Benton, M. J., 1994. Late Triassic to Middle Jurassic extinctions among continental tetrapods: Testing the pattern, in: Fraser, N. C., Sues, H-D., (Eds.), In the Shadow of the Dinosaurs. Cambridge University Press, Cambridge, pp. 366-397.
[15]
Bernardi, M., Petti, F. M., Porchetti, S. D., Avanzini, M., 2013. Large tridactyl footprints associated with a diverse ichnofauna from the Carnian of the Southern Alps. New Mexico Museum of Natural History and Science Bulletin, 61, 48-54.
[16]
Berner, R. A., Beerling, D. J., 2007. Volcanic degassing necessary to produce CaCO3 undersaturated ocean at the Triassic-Jurassic boundary. Palaeogeography, Palaeoclimatology, Palaeoecology, 244, 368-373.
[17]
Blackburn, T J., Olsen, P. E., Bowring, S. A., McLean, N. M., Kent, D. V., Puffer, J., McHone, G., Rasbury, E. T., Et-Touhami, M., 2013. Zircon U-Pb geochronology links the end-Triassic extinction with the Central Atlantic Magmatic Province. Science, 340, 941-945.
[18]
Bonis, N. R., Kürschner, W. M., 2012. Vegetation history, diversity patterns, and climate change across the Triassic/Jurassic boundary. Paleobiology, 38, 240-264.
[19]
Bonis, N. R., Kürschner, W., Krystyn, L., 2009. A detailed palynological study of the Triassic-Jurassic transition in key sections of the Eiberg basin (Northern calcareous Alps, Austria). Review of Palaeobotany and Palynology, 156, 376-400.
[20]
Brugman, W. A., 1983. Permian-Triassic Palynology. State University Utrecht, Utrecht, pp. 1-121.
[21]
Burgoyne, P. M., van Wyk, A. E., Anderson, J. M., Schrire, B. D., 2005. Phanerozoic evolution of plants on the African plate. Journal of African Earth Sciences, 43, 13-52.
[22]
Cascales-Miñana, B., Cleal, C. J., 2012. Plant fossil record and survival analysis. Lethaia, 45, 71-82.
[23]
Cirilli, S., 2010. Uppermost Triassic-lowermost Jurassic palynology and palynostratigraphy: A review, in: Lucas, S. G., (Ed.), The Triassic timescale. Geological Society, London, Special Publications, 334, 285-314.
[24]
Cirilli, S., Marzoli, A., Tanner, L. H., Bertrand, H., Buratti, N., Jourdan, F., Bellieni, G., Kontak, D., Renne, P. R., 2009. Late Triassic onset of the Central Atlantic Magmatic Province (CAMP) volcanism in the Fundy Basin (Nova Scotia): New stratigraphic constraints. Earth and Planetary Science Letters, 286, 514-525.
[25]
Cleal, C. J., 1993a. Pteridophyta, in: Benton, M. J., (Ed.), The Fossil Record 2. Chapman and Hall, London, pp. 779-794.
[26]
Cleal, C. J., 1993b. Gymnospermophyta, in: Benton, M. J., (Ed.), The Fossil Record 2. Chapman and Hall, London, pp. 795-808.
[27]
Cleveland, D. M., Nordt, L. C., Dworkin, S. I., Atchley, S. C., 2008. Pedogenic carbonate isotopes as evidence for extreme climatic events preceding the Triassic-Jurassic boundary: Implications for the biotic crisis? GSA Bulletin, 120, 1408-1415.
[28]
Colbert, E. H., 1949. Progressive adaptations as seen in the fossil record, in: Jepsen, G. L., Mayr, E., Simpson, G. G., (Eds.), Genetics, Paleontology and Evolution. Princeton University Press, Princeton, pp. 390-402.
[29]
Colbert, E. H., 1958. Triassic tetrapod extinction at the end of the Triassic Period. Proceedings National Academy of Science USA, 44, 973-977.
[30]
Cornet, B., 1977. The Palynostratigraphy and Age of the Newark Supergroup. Ph.D. Thesis. Pennsylvania State University, University Park, PA, 1-505.
[31]
Cornet, B., Olsen, P. E., 1985. A summary of the biostratigraphy of the Newark Supergroup of eastern North America with comments on provinciality, in: Weber, R., (Ed.), III Congreso Latinoamericano de Paleontologia Mexico, Simposio Sobre Floras del Triasico Tardio, su Fitogeografia y Paleoecologia, UNAM Instituto de Geologia, Mexico City, pp. 67-81.
[32]
Dam, G., Surlyk, F., 1993. Cyclic sedimentation in a large wave- and storm-dominated anoxic lake, Kap Stewart Formation (Rhaetian-Sinemurian), Jameson Land, East Greenland. International Association of Sedimentologists Special Publication, 18, 419-448.
[33]
Deckart, K., Fèraud, G., Bertrand, H., 1997. Age of Jurassic continental tholeiites of French Guyana, Surinam and Guinea: Implications for the initial opening of the Central Atlantic Ocean. Earth and Planetary Science Letters, 150, 205-220.
[34]
Desojo, J. B., Heckert, A. B., Martz, J. W., Parker, W. G., Schoch, R. R., Small, B. J., Sulej, T., 2013. Aetosauria: A clade of armoured pseudosuchians from the Upper Triassic continental beds, in: Nesbitt, S. J., Desojo, J. B., Irmis, R. B., (Eds.), Anatomy, phylogeny and palaeobiology of early archosaurs and their Kin. Geological Society, London, Special Publications, 379, 203-240.
[35]
Dzik, J., Sulej, T., Niedzwiedzki, G., 2008. A dicynodont-theropod association in the latest Triassic of Poland. Acta Palaeontologica Polonica, 53, 733-738.
[36]
Edwards, D., 1993. Bryophyta, in: Benton, M. J., (Ed.), The Fossil Record 2. Chapman and Hall, London, pp. 775-778.
[37]
Fedak, T. J., Sues, H.-D., Olsen, P. E., 2015. First record of the tritylodontid cynodont Oligokyphus and cynodont postcranial bones from the McCoy Brook Formation of Nova Scotia, Canada. Canadian Journal of Earth Sciences, 52, 244-249.
[38]
Fisher, M. J., Dunay, R. E., 1981. Palynology and the Triassic/Jurassic boundary. Review of Palaeobotany and Palynology, 34, 129-135.
[39]
Flügel, E., 2002. Triassic reef patterns, in: Kiessling, W., Flügel, E., Golonka, J., (Eds.), Phanerozoic reef patterns, SEPM Special Publication, 72, 391-463.
[40]
Fowell, S. J., Olsen, P. E., 1993. Time calibration of Triassic-Jurassic microfloral turnover, eastern North America. Tectonophysics, 222, 361-369.
[41]
Fowell, S. J., Traverse, A., 1995. Palynology and age of the upper Blomidon Formation, Fundy basin, Nova Scotia. Review of Palaeobotany and Palynology, 86, 211-233.
[42]
Fowell, S. J., Cornet, B., Olsen, P. E., 1994. Geologically rapid Late Triassic extinctions: palynological evidence from the Newark Supergroup. Geological Society of America Special Paper, 288, 197-206.
[43]
Fraser, N. C., 1994. Assemblages of small tetrapods from British Late Triassic fissure deposits, in: Fraser, N. C., Sues, H-D., (Eds.), In the Shadow of the Dinosaurs. Cambridge University Press, Cambridge, pp. 214-226.
[44]
Galli, M. T., Jadoul, F., Bernasconi, S. M., Weissert, H., 2005. Anomalies in global carbon cycling and extinction at the Triassic/Jurassic boundary: Evidence from a marine C-isotope record. Palaeogeography, Palaeoclimatology, Palaeoecology, 16, 203-214.
[45]
Giordano, N., Rigo, M., Ciarapica, G., Bertinelli, A., 2010. New biostratigraphical constraints for the Norian/Rhaetian boundary: Data from Lagonegro basin, Southern Appenines, Italy. Lethaia, 43, 573-586.
[46]
Gómez, J. J., Goy, A., Barrón, E., 2007. Events around the Triassic-Jurassic boundary in northern and eastern Spain: A review. Palaeogeography, Palaeoclimatology, Palaeoecology, 244, 89-110.
[47]
Grimaldi, D., Engel, M. S., 2005. Evolution of the Insects. Cambridge University Press, Cambridge, pp. 1-755.
[48]
Guex, J., Bartolin, A., Atudorei, V., Taylor, D., 2004. High-resolution ammonite and carbon isotope stratigraphy across the Triassic-Jurassic boundary at New York Canyon (Nevada). Earth and Planetary Science Letters, 225, 29-41.
[49]
Hallam, A., 1995. Major bio-events in the Triassic and Jurassic, in: Walliser, O., (Ed.), Global Events and Event Stratigraphy. Springer, Berlin, pp. 265-283.
[50]
Hallam, A., 2002. How catastrophic was the end-Triassic mass extinction? Lethaia, 35, 147-157.
[51]
Hallam, A., Wignall, P. B., 1997. Mass Extinctions and Their Aftermath. Oxford University Press, Oxford, pp.1-320.
[52]
Harris, T. M., 1937. The fossil flora of Scoresby Sound East Greenland, Part 5: Stratigraphic relations of the plant beds. Meddelelser om Grønland, 112, 1-112.
[53]
Haworth, M., Raschi, A., 2014. An assessment of the use of epidermal micro-morphological features to estimate leaf economics of Late Triassic-Early Jurassic fossil Ginkgoales. Review of Palaeobotany and Palynology, 205, 1-8.
[54]
Hesselbo, S. P., Robinson, S. A., Surlyk, F., Piasecki, S., 2002. Terrestrial and marine extinction at the Triassic-Jurassic boundary synchronized with major carbon-cycle perturbation: A link to initiation of massive volcanism? Geology, 30, 251-254. 2.0.CO;2 target="_blank">
[55]
Hillebrandt, A. v., Krystyn, L., Kürschner, W. M., Bonis, N. R., Ruhl, M., Richoz, S., Schobben, M. A. N., Ulrichs, M., Bown, P. R., Kment, K., McRoberts, C. A., Simms, M., Tomãsovych, A., 2013. The global stratotype section and point (GSSP) for the base of the Jurassic System at Kuhjoch (Karwendel Mountains, Northern Calcareous Alps, Tyrol, Austria). Episodes, 36, 162-198.
[56]
Hodych, J. P., Dunning, G. R., 1992. Did the Manicouaga impact trigger end-of-Triassic mass extinction? Geology, 20, 51-54. 2.3.CO;2 target="_blank">
[57]
Hornung, T., Brandner, R., Krystyn, L., Jaochimski, M. M., Keim, L., 2007. Multistratigraphic constrainst on the NW Tethyan "Carnian crisis". New Mexico Museum of Natural History and Science Bulletin, 41: 59-67.
[58]
Hounslow, M. W., Muttoni, G., 2010. The geomagnetic polarity timescale for the Triassic: Linkage to stage boundary definitions, in: Lucas, S. G., (Ed.), The Triassic timescale. Geological Society, London, Special Publications, 334, 61-102.
[59]
Hounslow, M. W., Posen, P. E., Warrington, G., 2004. Magnetostratigraphy and biostratigraphy of the Upper Triassic and lowermost Jurassic succession, St. Audrie’s Bay, UK. Palaeogeography, Palaeoclimatology, Palaeoecology, 213, 331-358.
[60]
Huber, P., Lucas, S. G., Hunt, A. P., 1993. Vertebrate biochronology of the Newark Supergroup Triassic, eastern North America. New Mexico Museum of Natural History and Science Bulletin, 3, 179-186.
[61]
Hunt, A. P., 1993. A revision of the Metoposauridae (Amphibia: Temnospondyli) of the Late Triassic with description of a new genus from the western United States. Museum of Northern Arizona Bulletin, 59, 67-97.
[62]
Hunt, A. P., Lucas, S. G., 1991. A new rhynchosaur from West Texas (USA) and the biochronology of Late Triassic rhynchosaurs. Palaeontology, 34, 191-198.
[63]
Johnson, L. A., Simms, M. J., 1989. The timing and cause of Late Triassic marine invertebrate extinctions: Evidence from scallops and crinoids, in: Donovan, K., (Ed.), Mass Extinctions: Processes and Evidence, Columbia University Press, New York, pp. 174-194.
[64]
Kelber, K.-P., 1998. Phytostratigraphische aspekte der makrofloren des süddeutschen Keupers. Documenta Naturae, 117, 89-115.
[65]
Kiessling, W., Aberhan, M., Brenneis, B., Wagner, P. J., 2007. Extinction trajectories of benthic organisms across the Triassic-Jurassic boundary. Palaeogeography, Palaeoclimatology, Palaeoecology, 244, 201-222.
[66]
Klein, H., Lucas, S. G., 2010. The Triassic footprint record of crocodylomorphs - A critical re-evaluation. New Mexico Museum of Natural History and Science Bulletin, 51, 55-60.
[67]
Klein, H., Lucas, S. G., 2013. The Late Triassic tetrapod ichnotaxon Apatopus lineatus (Bock 1952) and its distribution. New Mexico Museum of Natural History and Science Bulletin, 61, 313-324.
[68]
Knoll, A. H., 1984. Patterns of extinction in the fossil record of vascular plants, in: Nitecki, M., (Ed.), Extinction. University of Chicago Press, Chicago, pp. 21-68.
[69]
Kozur, H. W., Bachmann, G. H., 2005. Correlation of the Germanic Triassic with the international scale. Albertiana, 32, 21-35.
[70]
Kozur, H. W., Bachmann, G. H., 2008. Updated correlation of the Germanic Triassic with the Tethyan scale and assigned numeric ages. Berichte Geologische Bundesanstalt, 76, 53-58.
[71]
Kozur, H. W., Weems, R. E., 2005. Conchostracan evidence for a late Rhaetian to early Hettangian age for the CAMP volcanic event in the Newark Supergroup, and a Sevatian (late Norian) age for the immediately underlying beds. Hallesches Jahrbuch Geowissenschaft, B27, 21-51.
[72]
Kozur, H. W., Weems, R. E., 2007. Upper Triassic conchostracan biostratigraphy of the continental rift basins of eastern North America: Its importance for correlating Newark Supergroup events with the Germanic Basin and the international geologic time scale. New Mexico Museum of Natural History and Science Bulletin, 41, 137-188.
[73]
Kozur, H. W., Weems, R. E., 2010. The biostratigraphic importance of conchostracans in the continental Triassic of the northern hemisphere, in: Lucas, S. G., (Ed.), The Triassic timescale. Geological Society, London, Special Publications, 334, 315-417.
[74]
Krystyn, L., Boquerel, H., Kuerschner, W., Richoz, S., Gallet, Y., 2007. Proposal for a candidate GSSP for the base of the Rhaetian Stage. New Mexico Museum of Natural History and Science Bulletin, 41, 189-199.
[75]
Kuerschner, W. M., Bonis, N. R., Krystyn, L., 2007. Carbon-isotope stratigraphy and palynostratigraphy of the Triassic-Jurassic transition in the Tiefengraben section - Northern Calcareous Alps (Austria). Palaeogeography, Palaeoclimatology, Palaeoecology, 244, 257-280.
[76]
Kürschner, W. M., Herngreen, G. F. W., 2010. Triassic palynology of central and northwestern Europe: A review of palynofloral diversity patterns and biostratigraphic subdivisions, in: Lucas, S. G., (Ed.), The Triassic timescale. Geological Society, London, Special Publications, 334, 263-283.
[77]
Labandeira, C. C., Sepkoski, J. J., Jr., 1993. Insect diversity in the fossil record. Science, 261, 310-315.
[78]
Langer, M. C., 2014. The origins of Dinosauria: Much ado about nothing. Palaeontology, 2014, 1-10.
[79]
Langer, M. C., Ezcurra, M. D., Bittencourt, J. S., Novas, F. E., 2009. The origin and early evolution of dinosaurs. Biological Reviews, 84, 1-56.
[80]
Long, R. A., Murry, P. A., 1995, Late Triassic (Carnian and Norian) tetrapods from the southwestern United States. New Mexico Museum of Natural History and Science Bulletin, 4, 1-254.
[81]
Lucas, S. G., 1994. Triassic tetrapod extinctions and the compiled correlation effect. Canadian Society of Petroleum Geologists Memoir, 17, 869-875.
[82]
Lucas, S. G., 1998. Global Triassic tetrapod biostratigraphy and biochronology. Palaeogeography, Palaeoclimatology, Palaeoecology, 143, 347-384.
[83]
Lucas, S. G., 2007. Tetrapod footprint biostratigraphy and biochronology. Ichnos, 14, 5-38.
[84]
Lucas, S. G., 2008. Global Jurassic tetrapod biochronology. Volumina Jurassica, 6, 99-108.
[85]
Lucas, S. G., 2010a. The Triassic timescale: An introduction, in: Lucas, S. G., (Ed.), The Triassic timescale. Geological Society, London, Special Publications, 334, 1-16.
[86]
Lucas, S. G., 2010b. The Triassic timescale based on nonmarine tetrapod biostratigraphy and biochronology, in: Lucas, S. G., (Ed.), The Triassic timescale. Geological Society, London, Special Publications, 334, 447-500.
[87]
Lucas, S. G., 2013a. Plant megafossil biostratigraphy and biochronology, Upper Triassic Chinle Group, western USA. New Mexico Museum of Natural History and Science Bulletin, 61, 354-365.
[88]
Lucas, S. G., 2013b. A new Triassic timescale. New Mexico Museum of Natural History and Science Bulletin, 61, 366-374.
[89]
Lucas, S. G., 2015. Age and correlation of Late Triassic tetrapods from southern Poland. Annales Societatis Geologorum Poloniae, in press. doi.org/10.14241/asgp.2015.024.
[90]
Lucas, S. G., Heckert, A. B., 2011. Late Triassic aetosaurs as the trackmaker of the tetrapod footprint ichnotaxon Brachychirotherium. Ichnos, 18, 197-208.
[91]
Lucas, S. G., Huber, P., 2003. Vertebrate biostratigraphy and biochronology of the nonmarine Late Triassic, in: LeTourneau, P. M., Olsen, P. E., (Eds.), The great rift valleys of Pangea in eastern North America. Volume 2. Sedimentology, Stratigraphy, and Paleontology. Columbia University Press, New York, pp. 143-191.
[92]
Lucas, S. G., Hunt, A. P., 1994. The chronology and paleobiogeography of mammalian origins, in: Fraser, N. C., Sues, H-D.,(Eds.), In the Shadow of the Dinosaurs. Cambridge University Press, Cambridge, pp. 335-351.
[93]
Lucas, S. G., Tanner, L. H., 2004. Late Triassic extinction events. Albertiana, 31, 31-40.
[94]
Lucas, S. G., Tanner, L. H., 2007a. Tetrapod biostratigraphy and biochronology of the Triassic-Jurassic transition on the southern Colorado Plateau, USA. Palaeogeography, Palaeoclimatology, Palaeoecology, 244, 242-256.
[95]
Lucas, S. G., Tanner, L. H., 2007b. The nonmarine Triassic-Jurassic boundary in the Newark Supergroup of eastern North America. Earth Science Reviews, 84, 1-20.
[96]
Lucas, S. G., Tanner, L. H., 2008. Reexamination of the end-Triassic mass extinction, in: Elewa, A. M. T., (Ed.), Mass Extinction. Springer Verlag, New York, pp. 66-103.
[97]
Lucas, S. G., Wild, R., 1995. A Middle Triassic dicynodont from Germany and the biochronology of Triassic dicynodonts. Stuttgarter Beiträge zur Naturkunde, 220, 1-16.
[98]
Lucas, S. G., Heckert, A. B., Hotton, N. III., 2002. The rhynchosaur Hyperodapedon from the Upper Triassic of Wyoming and its global biochronological significance. New Mexico Museum of Natural History and Science Bulletin, 21, 149-156.
[99]
Lucas, S. G., Lockley, M. G., Hunt, A. P., Milner, A. R. C., Tanner, L. H., 2006a. Tetrapod footprint biostratigraphy of the Triassic-Jurassic transition in the American Southwest. New Mexico Museum of Natural History and Science Bulletin, 37, 105-108.
[100]. Lucas, S. G., Klein, H., Lockley, M. G., Spielmann, J. A., Gierlinski, G., Hunt, A. P., Tanner, L. H., 2006b. Triassic-Jurassic stratigraphic distribution of the theropod footprint ichnogenus Eubrontes. New Mexico Museum of Natural History and Science Bulletin, 37, 86-93.
[101]. Lucas, S. G., Hunt, A. P., Heckert, A. B., Spielmann, J. A., 2007a. Global Triassic tetrapod biostratigraphy and biochronology: 2007 status. New Mexico Museum of Natural History and Science Bulletin, 41, 229-240.
[102]. Lucas, S. G., Taylor, D. G., Guex, J., Tanner, L. H., Krainer, K., 2007b. The proposed global stratotype section and point for the base of the Jurassic System in the New York Canyon area, Nevada, USA. New Mexico Museum of Natural History and Science Bulletin, 40, 139-168.
[103]. Lucas, S. G., Tanner, L. H., Donohoo-Hurley, L. L., Geissman, J. W., Kozur, H. W., Heckert, A. B., Weems, R. E., 2011. Position of the Triassic-Jurassic boundary and timing of the end-Triassic extinctions on land: Data from the Moenave Formation on the southern Colorado Plateau, USA. Palaeogeography, Palaeoecology, Palaeoclimatology, 302, 194-205.
[104]. Lucas, S. G., Tanner, L. H., Kozur, H. W., Weems, R. E., Heckert, A. B., 2012. The late Triassic timescale: Age and correlation of the Carnian-Norian boundary. Earth Science Reviews, 114, 1-8.
[105]. Lucas, S. G., Szajna, M. J., Lockley, M. G., Fillmore, D. L., Simpson, E. L., Klein, H. Boyland, J., Hartline, B. W., 2014. The Middle-Late Triassic tetrapod footprint ichnogenus Gwynnedichnium. New Mexico Museum of Natural History and Science Bulletin, 62, 135-156.
[106]. Maisch, M. W., Kapitzke, M., 2010. A presumably marine phytosaur (Reptilia: Archosauria) from the pre-planorbis beds (Hettangian) of England. Neues Jahrbuch für Geologie und Paläontologie Abhandlungen, 257, 373-379.
[107]. Mander, L., Kürschner, W. M., McElwain, J. C., 2010. An explanation for conflicting records of Triassic-Jurassic plant diversity. Proceedings of the National Academy of Science USA, 107, 15351-15356.
[108]. Mander, L., Wesseln, C. J., McElwain, J. C., Punyasena, S. W., 2012. Tracking taphonomic regimes using chemical and mechanical damage of pollen and spores: An example from the Triassic-Jurassic mass extinction. PLOS ONE, 7(11), e49153.
[109]. Mander, L., Kürschner, W. M., McElwain, J. C., 2013. Palynostratigraphy and vegetation history of the Triassic-Jurassic transition in East Greenland. Journal of the Geological Society London, 170, 37-46.
[110]. Marzoli, A., Renne, P. R., Piccirillo, E. M., Ernesto, M., Bellieni, G., DeMin, A., 1999. Extensive 200-million-year-old continental flood basalts of the central Atlantic Magmatic province. Science, 284, 616-618.
[111]. Marzoli, A., Bertrand, H., Knight, K. B., Cirilli, S., Buratti, N., Verati, C., Nomade, S., Renne, P. R., Youbi, N., Martini, R., Allenbach, N., Neuwerth, R., Rapaille, C., Zaninetti, L., Zaninetti, L., Bellieni, G., 2004. Synchrony of the Central Atlantic magmatic province and the Triassic-Jurassic boundary climate and biotic crisis. GSA Bulletin, 32, 973-976.
[112]. Marzoli, A., Jourdan, F. Puffer, J. H., Cuppone, T., Tanner, L. H., Weems, R. E., Bertrand, H., Cirilli, S., Bellieni, G., De Min, A., 2011. Timing and duration of the Central Atlantic magmatic province in the Newark and Culpeper basins, eastern U.S.A. Lithos, 122, 175-188.
[113]. McCune, A. R., Schaeffer, B., 1986. Triassic and Jurassic fishes: Patterns of diversity, in: Padian, K., (Ed.), The Beginning of the Age of Dinosaurs. Cambridge University Press, Cambridge, pp. 171-181.
[114]. McElwain, J. C., Punyasena, S. W., 2007. Mass extinction events and the plant fossil record. Trends in Ecology and Evolution, 22, 548-557.
[115]. McElwain, J. C., Beerling, D. J., Woodward, F. I., 1999. Fossil plants and global warming at the Triassic-Jurassic boundary. Science, 285, 1386-1390.
[116]. McElwain, J. C., Popa, M. E., Hesselbo, S. P., Haworth, M., Surlyk, F., 2007. Macroecological responses of terrestrial vegetation to climatic and atmospheric change across the Triassic/Jurassic boundary in East Greenland. Paleobiology, 33, 547-573.
[117]. McElwain , J. C., Wagner, P. J., Hesselbo, S. P., 2009. Fossil plant relative abundances indicate sudden loss of late Triassic biodiversity in East Greenland. Science, 324, 1554-1556.
[118]. McGhee, G. R., Jr., Sheehan, P. M., Bottjer, D. J., Droser, M. L., 2004. Ecological ranking of Phanerozoic biodiversity crises: Ecological and taxonomic severities are decoupled. Palaeogeography, Palaeoclimatology, Palaeoecology, 211, 289-297.
[119]. McGhee, G. R., Jr., Clapham, M. E., Sheehan, P. M., Bottjer, D. J., Droser, M. L., 2013. A new ecological-severity ranking of major Phanerozoic biodiversity crises. Palaeogeography, Palaeoclimatology, Palaeoecology, 370, 260-270.
[120]. McHone, J. G., Puffer, J. H., 1996. Hettangian flood basalts across the Pangaean rift. Connecticut State Geological and Natural History Survey Natural Resources Center Miscellaneous Report, 1, 29.
[121]. Milner, A. R., 1993. Amphibian-grade Tetrapoda, in: Benton, M. J., (Ed.), The Fossil Record 2. Chapman and Hall, London, pp. 665-679.
[122]. Milner, A. R., 1994. Late Triassic and Jurassic amphibians: Fossil record and phylogeny, in: Fraser, N. C., Sues, H-D., (Eds.), In the Shadow of the Dinosaurs. Cambridge University Press, Cambridge, pp. 5-22.
[123]. Milner, A. R. C., Kirkland, J. I., Birthisel, T. A., 2006. The geographic distribution and biostratigraphy of Late Triassic-Early Jurassic freshwater fish faunas of the western United States. New Mexico Museum of Natural History and Science Bulletin, 37, 522-529.
[124]. Nesbitt, S. J., 2011. The early evolution of archosaurs: Relationships and the origin of major clades. Bulletin of the American Museum of Natural History, 352, 1-292.
[125]. Nesbitt, S. J., Brusatte, S. L., Desojo, J. B., Liparini, A., De Franca, M. A. G., Weinbaum, J. C., Gower, D. J., 2013. Rauisuchia, in: Nesbitt, S. J., Desojo, J. B., Irmis, R. B., (Eds.), Anatomy, phylogeny and palaeobiology of early archosaurs and their kin. Geological Society, London, Special Publications, 379, 241-274.
[126]. Newham, E., Benson, R., Upchurch, P., Goswani, A., 2014. Mesozoic mammaliaform diversity: The effect of sampling corrections on reconstructions of evolutionary dynamics. Palaeogeography, Palaeoclimatology, Palaeoecology, 412, 32-44.
[127]. Nied?wiedzki, G., 2011. A Late Triassic dinosaur-dominated ichnofauna from the Tomanová Formation of the Tatra Mountains, central Europe. Acta Palaeontologica Polonica, 56, 291-300.
[128]. Niklas, K. J., Tiffney, B. H., Knoll, A. H., 1983. Patterns in vascular land plant diversification: A statistical analysis at the species level. Nature, 303, 614-616.
[129]. Ogg, J. G., 2012a. Triassic. In: Gradstein, F. M., Ogg, J. G., Schmitz, M. D., Ogg, G. M., (Eds.), The Geologic Timescale 2012. volume 2. Elsevier, Amsterdam, pp. 681-730.
[130]. Ogg, J. G., 2012b. Jurassic, in: Gradstein, F. M., Ogg, J. G., Schmitz, M. D., Ogg, G. M., (Eds.), The Geologic Timescale 2012. volume 2. Elsevier, Amsterdam, pp. 731-791.
[131]. Ogg, J. G., Huang, C., Hinnov, L., 2014. Triassic timescale status: A brief overview. Albertiana, 41, 3-30.
[132]. Olsen, P. E., Galton, P. M., 1984. A review of the reptile and amphibian assemblages from the Stormberg of South Africa, with special emphasis on the footprints and the age of the Stormberg. Palaeontologica Africana, 25, 87-110.
[133]. Olsen, P. E., Rainforth, E. C., 2003. The Early Jurassic ornithischian dinosaur ichnogenus Anomoepus, in: LeTourneau, P.M., Olsen, P. E., (Eds.), The Great Rift Valleys of Pangea in Eastern North America. volume 2. Sedimentology, Stratigraphy, and Paleontology. Columbia University Press, New York, pp. 314-368.
[134]. Olsen, P. E., Sues, H-D., 1986. Correlation of continental Late Triassic and Early Jurassic sediments, and patterns of the Triassic-Jurassic tetrapod transition, in: Padian, K., (Ed.), The Beginning of the Age of Dinosaurs. Cambridge University Press, Cambridge, pp. 321-351.
[135]. Olsen, P. E., Shubin, N. H., Anders, M. H., 1987. New Early Jurassic tetrapod assemblages constrain Triassic-Jurassic tetrapod extinction event. Science, 237, 1025-1029.
[136]. Olsen, P. E., Fowell, S. J., Cornet, B., 1990. The Triassic/Jurassic boundary in continental rocks of eastern North America; a progress report. Geological Society of America Special Paper, 247, 585-593.
[137]. Olsen, P. E., Schlische, R. W., Fedosh, M. S., 1996. 580 ky duration of the Early Jurassic flood basalt event in eastern North America estimated using Milankovitch cyclostratigraphy. Museum of Northern Arizona Bulletin, 60, 11-22.
[138]. Olsen, P. E., Smith, J. B., McDonald, N. G., 1998. Type material of the type species of the classic theropod footprint genera Eubrontes, Anchisauripus, and Grallator (Early Jurassic, Hartford and Deerfield basins, Connecticut and Massachusetts, U.S.A.). Journal of Vertebrate Paleontology, 18, 586-601.
[139]. Olsen, P. E., Kent, D. V., Sues, H. D., Koeberl, C., Huber, H., Montanari, A., Rainforth, E. C., Powell, S. J., Szajna, M. J., Hartline, B. W., 2002a. Ascent of dinosaurs linked to an iridium anomaly at the Triassic-Jurassic boundary. Science, 296, 1305-1307.
[140]. Olsen, P. E., Koeberl, C., Huber, H., Montanari, A., Fowell, S. J., Et-Touhani, M., Kent, D. V., 2002b. The continental Triassic-Jurassic boundary in central Pangea: Recent progress and preliminary report of an Ir anomaly. Geological Society of America Special Paper, 356, 505-522.
[141]. Olsen, P. E., Kent, D. V., Whiteside, J. H., 2011. Implications of the Newark Supergroup-based astrochronology and geomagnetic polarity time scale (Newark-APTS) for the tempo and mode of the early diversification of the Dinosauria. Earth and Environmental Science Transactions of the Royal Society of Edinburgh, 101, 201-229.
[142]. Olsen, P. E., Reid, J. C., Taylor, K. B., Whiteside, J. H., Kent, D. V., 2015. Revised stratigraphy of Late Triassic age strata of the Dan River basin (Virginia and North Carolina, USA) based on drill core and outcrop data. Southeastern Geology, 51, 1-31.
[143]. Onoue, T., Sato, H., Nakamura, T., Noguchi, T., Hidaka, Y., Shirai, N., Ebihara, M., Osawa, T., Hatsukawa, Y., Toh, Y., Koizumi, M., Harada, H., Orchard, M. J., Nedachi, M., 2012. Deep-sea record of impact apparently unrelated to mass extinction in the Late Triassic. Proceedings of the National Academy of Science, 109, 19134-19139.
[144]. Orbell, G., 1973. Palynology of the British Rhaeto-Liassic. Bulletin of the Geological Society of Great Britain, 44, 1-44.
[145]. Orchard, M. J., 2010. Triassic conodonts and their role in stage boundary definition, in: Lucas, S. G., (Ed.), The Triassic timescale. Geological Society, London, Special Publications, 334, 139-161.
[146]. Orchard, M. J., 2013. Five new genera of conodonts from the Carnian-Norian boundary beds of Black Bear Ridge, northeast British Columbia, Canada. New Mexico Museum of Natural History and Science Bulletin, 61, 445-457.
[147]. Orchard, M. J., 2014.Conodonts from the Carnian-Norian boundary (Upper Triassic) of Black Bear Ridge, northeastern British Columbia, Canada. New Mexico Museum of Natural History and Science Bulletin, 64, 1-139.
[148]. Pedersen, K. R., Lund, J. J., 1980. Palynology of the plant-bearing Rhaetian to Hettangian Kap Stewart Formation, Scoresby Sund, East Greenland. Review of Palaeobotany and Palynology, 31, 1-69.
[149]. Pie��kowski, G., Nied?wiedzki, G., Branski, P., 2014. Climatic reversals related to the Central Atlantic magmatic province caued the end-Triassic biotic crisis - Evidence from continental strata in Poland. Geological Society of America Special Paper, 505, 263-286.
[150]. Pie��kowski, G., Nied?wiedzki, G., Waksmundzka, M., 2011. Sedimentological, palynological and geochemical studies of the terrestrial Triassic-Jurassic boundary in northwestern Poland. Geological Magazine, 149, 308-332.
[151]. Pott, C., McLoughlin, S., 2009. Bennettitalean foliage in the Rhaetian-Bajocian (latest Triassic-Middle Jurassic) floras of Scania, southern Sweden. Review of Palaeobotany and Palynology, 158, 117-166.
[152]. Rainforth, E. C., 2003. Revision and re-evaluation of the Early Jurassic dinosaurian ichnogenus Otozoum. Palaeontology, 46, 803-838.
[153]. Ramezani, J., Bowring, S. A., Pringle, M., Winslow III, F. D., Rasbury, E. T., 2005. The Manicouagan impact melt rock: A proposed standard for intercalibration of U-Pb and 40Ar/39/Ar isotopic systems. 15th V.M. Goldschmidt Conference Abstract Volume, A321.
[154]. Rigo, M., Preto, N., Roghi, G., Tateo, F., Mietto, P., 2007. A rise in the carbonate compensation depth of western Tethys in the Carnian (Late Triassic): Deep-water evidence for the Carnian pluvial event. Palaeogeography, Palaeoclimatology, Palaeoecology, 246, 188-205.
[155]. Ruckwied, K., Götz, A. E., Pálfy, J., Török, Á., 2008. Palynology of a terrestrial coal-bearing series across the Triassic/Jurassic boundary (Mecsek Mts, Hungary). Central European Geology, 51, 1-15.
[156]. Ruhl, M., Veld, H., Kuerschner, W. M., 2010. Sedimentary organic matter characterization of the Triassic-Jurassic boundary GSSP at Kuhjoch (Austria). Earth and Planetary Science Letters, 292, 17-26.
[157]. Schäfer, P., Fois, E., 1987. Systematics and evolution of Triassic Bryozoa. Geologica et Palaeontologica, 21, 173-225.
[158]. Schaller, E. M., Wright, J. D., Kent, D. V., Olsen, P. E., 2012. Rapid emplacement of the Central Atlantic Magmatic Province as net sink for CO2. Earth and Planetary Science Letters, 323-324, 27-39.
[159]. Schoch, R. R., Milner, A. R., 2000. Stereospondyli. Encyclopedia of Paleoherpetology, 3B, 1-203.
[160]. Schoene, B., Guex, J., Bartolini, A., Schaltegger, U., Blackburn, T. J., 2010, Correlating the end-Triassic mass extinction and flood basalt volcanism at the 100 ka level. Geology, 38, 387-390.
[161]. van de Schootbrugge, B., Quan, T., Lindström, S., Püttmann, W., Heunisch, C., Pross, J., Fiebig, J., Petschick, R., Röhling, H-G., Richoz, S., Rosenthal, Y., Falkowski, P. G., 2009. Floral changes across the Triassic/Jurassic boundary linked to flood basalt volcanism. Nature Geoscience, 2, 589-594.
[162]. Schuurman, W. M. L., 1979. Aspects of Late Triassic palynology. 3. Palynology of latest Triassic and earliest Jurassic deposits of the northern limestone Alps in Austria and southern Germany, with special reference to a palynological characterization of the Rhaetian stage in Europe. Review of Palaeobotany and Palynology, 27, 53-75.
[163]. Sepkoski, J. J., Jr., 1982. Mass extinctions in the Phanerozoic oceans: A review. Geological Society of America Special Paper, 190, 283-289.
[164]. Sha, J., Vajda, V., Pan, Y., Larsson, L., Yao, X., Zhang, X., Wang, Y., Cheng, X., Jiang, B., Deng, S., Chen, S., Peng, B., 2011. Stratigraphy of the Triassic-Jurassic boundary successions of the southern margin of the Junggar basin, northwestern China. Acta Geologica Sinica, 85, 421-436.
[165]. Sha, J., Olsen, P. E., Pan, Y., Xu, D., Wang, Y., Zhang, X., Yao, X., Vajda, V., 2015. Triassic-Jurassic climate in continental high-latitude Asia was dominated by obliquity-paced variations (Junggar Basin, Ürümqi, China). Proceedings of the National Academy of Sciences, 112, 3624-3629.
[166]. Shubin, N. H., Olsen, P. E., Sues, H.-D., 1994. Early Jurassic small tetrapods from the McCoy Brook Formation of Nova Scotia, Canada, in: Fraser, N. C., Sues, H-D., (Eds.), In the Shadow of the Dinosaurs. Cambridge University Press, Cambridge, pp. 242-250.
[167]. Sigurdsson, H., 1990. Assessment of atmospheric impact of volcanic eruptions. Geological Society of America Special Paper, 247, 99-110.
[168]. Silvestri, S. M., Szajna, M. J., 1993. Biostratigraphy of vertebrate footprints in the Late Triassic section of the Newark Basin, Pennsylvania: Reassessment of stratigraphic ranges. New Mexico Museum of Natural History and Science Bulletin, 3, 439-445.
[169]. Simms, M. J., Ruffell, A. H., 1989. Synchroneity of climatic change and extinctions in the Late Triassic. Geology, 17, 265-268. 2.3.CO;2 target="_blank">
[170]. Simms, M. J., Ruffell, A. H., 1990. Climatic and biotic change in the Late Triassic. Journal Geological Society (London), 147, 321-327.
[171]. Smith, R. M. H., Marsicano, C. A., Wilson, J. A., 2009. Sedimentology and paleoecology of a diverse Early Jurassic tetrapod tracksite in Lesotho, southern Africa. Palaios, 24, 672-684.
[172]. Spielmann, J. A., Lucas, S. G., Hunt, A. P., 2013. The first Norian (Revueltian) rhynchosaur: Bull Canyon Formation, New Mexico, U.S.A. New Mexico Museum of Natural History and Science Bulletin, 61, 562-566.
[173]. Steinthorsdottir, M., Jeram, A. J., McElwain, J. C., 2011. Extremely elevated CO2 concentrations at the Triassic/Jurassic boundary. Palaeogeography, Palaeoclimatology, Palaeoecology, 308, 418-432.
[174]. Stocker, M. R., Butler, R. J., 2013. Phytosauria, in: Nesbitt, S.J., Desojo, J. B., Irmis, R. B., (Eds.), Anatomy, phylogeny and palaeobiology of early archosaurs and their kin. Geological Society, London, Special Publications, 379, 91-118.
[175]. Sues, H-D., Olsen, P. E., 2015. Stratigraphic and temporal context and faunal diversity of Permian-Jurassic continental tetrapod assemblages from the Fundy rift basin, eastern Canada. Atlantic Geology, 51, 139-205.
[176]. Szajna, M. J., Silvestri, S. M., 1996. A new occurrence of the ichnogenus Brachychirotherium: Implications for the Triassic-Jurassic mass extinction event. Museum of Northern Arizona Bulletin, 60, 275-283.
[177]. Szulc, J., Racki, G., ?rodo��, J., Lucas, S. G., 2015. Norian age of the Lipie ?l?skie vertebrate fauna explains its alleged oddity. Acta Palaeontologica Polonica, in press.
[178]. Tanner, L. H., Hubert, J. F., Coffey, B. P., McInerney, D. P., 2001. Stability of atmospheric CO2 levels across the Triassic/Jurassic boundary. Nature, 411, 675-677.
[179]. Tanner, L. H., Lucas, S. G., Chapman, M. G., 2004. Assessing the record and causes of Late Triassic extinctions. Earth-Science Reviews, 65, 103-139.
[180]. Tanner, L. H., Smith, D. L., Allan, A., 2007. Stomatal response of swordfern to volcanogenic CO2 and SO2 from Kilauea volcano, Hawaii. Geophysical Research Letters, 34, L15807,http://dx.doi.org/10.1029/2007GL030320.
[181]. Thulborn, T., Turner, S., 2003. The last dicynodont: An Australian Cretaceous relict. Proceedings Royal Society London B, 270, 985-993.
[182]. Traverse, A., 1988. Plant evolution dances to a different beat. Historical Biology, 1, 277-301.
[183]. de Valais, S., 2009. Ichnotaxonomic revision of Ameghinichnus, a mammalian ichnogenus from the Middle Jurassic La Matilde Formation, Santa Cruz Province, Argentina. Zootaxa, 2203, 1-21.
[184]. Visscher, H., Van Houte, M., Brugman, W. A., Poort, R. J., 1994. Rejection of a Carnian (late Triassic) "pluvial event" in Europe. Review of Palaeobotany and Palynology, 83, 217-226.
[185]. Weems, R. E., 1992. The "terminal Triassic catastrophic extinction event" in perspective: A review of Carboniferous through Early Jurassic terrestrial vertebrate extinction patterns. Palaeogeography, Palaeoclimatology, Palaeoecology, 94, 1-29.
[186]. Weems, R. E., Lucas, S. G., 2015. A revision of the Norian conchostracan zonation in North America and its implications for Late Triassic North American tectonic history. New Mexico Museum of Natural History and Science Bulletin, 67, 303-317.
[187]. Whiteside, J. H., Olsen, P. E., Kent, D. V., Fowell, S. J., Et-Touhami, M., 2007. Synchrony between the Central Atlantic magmatic province and the Triassic-Jurassic mass-extinction event? Palaeogeography, Palaeoclimatology, Palaeoecology, 244, 345-367.
[188]. Whiteside, J. H., Olsen, P. E., Eglinton, T., Brookfield, M. E., Sambrotto, R. N., 2010. Compound-specific carbon isotopes from Earth’s largest flood basalt eruptions directly linked to the end-Triassic mass extinction. Proceedings National Academy of Science USA, 107, 6721-6725.
[189]. Wotzlaw, J-F., Guex, J., Bartolini, A., Gallet, Y., Krystyn, L., McRoberts, C. A., Taylor, D., Schoene, B., Schaltegger, U., 2014. Towards accurate numerical calibration of the Late Triassic: High-precision U-Pb geochronology constraints on the duration of the Rhaetian. Geology, 42, 571-574.
[190]. Yapp, C. J., Poths, H., 1996. Carbon isotopes in continental weathering environments and variations in ancient atmospheric CO2 pressure. Earth and Planetary Science Letters, 137, 71-82.