Cretaceous dinosaur and crocodylomorph egg records of the Sanshui Basin, Guangdong Province, southern China

doi:10.1016/j.jop.2023.04.002

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Abstract

This paper describes newly discovered dinosaur and crocodylomorph egg fragments from the Upper Cretaceous Sanshui and Dalangshan formations of the Sanshui Basin of Guangdong Province, southern China. Despite the absence of macroscopic information, the eggshell specimens can be identified to the oofamilies Prismatoolithidae, Elongatoolithidae and Krokolithidae, and it is hypothesized that these three families of eggs correspond to troodontids, oviraptorids and crocodiloids, respectively. Comparison with egg fossils from Nanxiong Basin, Heyuan Basin and Ganzhou Basin highlights the similarity between these egg assemblages and those from Nanxiong Basin. The egg fossils found in the Sanshui and the Dalangshan formations enrich the known faunal types in the research area and facilitate further studies on the diversity of Late Cretaceous vertebrate assemblages in the Sanshui Basin.

Key words： Prismatoolithidae Elongatoolithidae Krokolithidae Sanshui Formation Dalangshan Formation Upper Cretaceous

Received: 16 November 2022

Corresponding Authors: ^*E-mail: xinglida@cugb.edu.cn (L.-D. Xing).

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http://journal09.magtechjournal.com/Jweb_jop/EN/10.1016/j.jop.2023.04.002 OR http://journal09.magtechjournal.com/Jweb_jop/EN/Y2023/V12/I3/401

[1] Andrews R.C.,1933. The new conquest of Central Asia: A narrative of the explorations of the Central Asiatic Expeditions in Mongolia and China, 1921-1930. Nature, 132(3324), 81-83. https://doi.org/10.1038/132081a0.
[2] Antunes M.T., Taquet P., Ribeiro V., 1998. Upper Jurassic dinosaur and crocodile nesting site, Lourinhã, Portugal.Memórias da Academia das Ciências de Lisboa, 37, 83-99.
[3] Bray E.S.,1996. Eggs and eggshells from the Upper Cretaceous North Horn Formation, central Utah.Vertebrate Paleontology in Utah, Utah Geological Survey Miscellaneous Publication, 99(1), 361-375.
[4] Chang Y.P., Tung Y.S., 1963. Subdivision of "redbeds" of Nanhsiung Basin, Kwangtung.Vertebrata PalAsiatica, 7(3), 249-260 (in Chinese with English abstract).
[5] Chen P.P.,2018. The Response of Late Cretaceous-Eocene Epoch Volcanic and Sedimentary Sequence in Sanshui Basin to the Tectonic Evolution of the Northern Margin of South China Sea (Ph.D. thesis). Beijing: China University of Geosciences (Beijing), pp. 12-14 (in Chinese).
[6] Cheng Y.N., Ji Q., Wu X.C., Shan H.Y., 2008. Oviraptorosaurian eggs (Dinosauria) with embryonic skeletons discovered for the first time in China. Acta Geologica Sinica (English Edition), 82(6), 1089-1094. https://doi.org/10.1111/j.1755-6724.2008.tb00708.x.
[7] Dong Z.M.,1979. Dinosaurs from the Cretaceous of South China. Institute of Vertebrate Paleontology and Paleoanthropology, Nanjing Institute of Paleontology, Science Press, pp. 342-350.
[8] Dong Z.M.,1992. Dinosaurian Faunas of China. Springer-Verlag, Berlin, pp. 1-188.
[9] Fang X.S., Li P.X., Zhang Z.J., Zhang X.Q., Lin Y.L., Guo S.B., Cheng Y.M., Li Z.Y., Zhang X.J., Cheng Z.W.,2009a. Cretaceous strata in Nanxiong Basin of Guangdong and the evolution from the dinosaur egg to the bird egg. Acta Geoscientica Sinica, 30(2), 167-186. https://doi.org/10.3321/j.issn:1006-3021.2009.02.004 (in Chinese with English abstract).
[10] Fang X.S., Wang Y.Z., Jiang Y.G., 2000. On the Late Cretaceous fossil eggs of Tianta, Zhejiang. Geological Review, 46(1), 105-112. https://doi.org/10.1007/s11769-000-0051-4 (in Chinese with English abstract).
[11] Fang X.S., Yue Z., Ling H.,2009b. Review of the past 15 years' research on fossil eggs in China. Acta Geoscientica Sinica, 30(4), 523-542. https://doi.org/10.3321/j.issn:1006-3021.2009.04.012 (in Chinese with English abstract).
[12] Fang X.S., Zhang Z.J., Pang Q.Q., Li P.X., Han Y.J., Xie H.L., Yan R.H., Pang F.J., Lu J.L., Cheng Z.W.,2007. Cretaceous strata and egg fossils in Xixia, Henan Province. Acta Geoscientica Sinica, 28(2), 123-142. https://doi.org/10.3321/j.issn:1006-3021.2007.02.003 (in Chinese with English abstract).
[13] Garcia G.,2000. New data on the fossil eggshell diversity from the lower Maastrichtian of southern France. Geobios, 33(1), 113-126. https://doi.org/10.1016/S0016-6995(00)80155-2.
[14] He F.L., Huang X.J., Li X.Y.,2017. Occurrence rule and buried characteristics of dinosaur fossils in the Ganzhou Basin, Jiangxi Province. East China Geology, 38(4), 250-254. https://doi.org/10.16788/j.hddz.32-1865/P.2017.04.002 (in Chinese with English abstract).
[15] Hirsch K.F.,1985. Fossil crocodilian eggs from the Eocene of Colorado.Journal of Vertebrate Paleontology, 59(3), 531-542.
[16] Hirsch K.F.,1994a. Upper Jurassic eggshells from the Western Interior of North America. In: Carpenter, K., Hirsch, K.F., Horner, J.R. (Eds.), Dinosaur Eggs and Babies. Cambridge: Cambridge University Press, pp. 137-150.
[17] Hirsch K.F.,1994b. The fossil record of vertebrate egg. In: Donovan, S.K. (Ed.), The Palaeobiology of Trace Fossils. John Wiley and Sons, London, pp. 269-294.
[18] Horner J.R., Weishampel D.B., 1996. Correction: A comparative embryological study of two ornithischian dinosaurs. Nature, 383(6595), 103. https://doi.org/10.1038/383103b0.
[19] Hou H.M., Yuan Y.R., Zhang Y.X., Tang X.Z., Wang B.G., 1994. Paleomagnetic study of Sanshui Basin in Guangdong from Late Cretaceous to Eocene and its tectonic significance.Tropic Oceanology, 13(4), 65-72 (in Chinese with English abstract).
[20] Hou M.C., Chen H.D., Li Z.Z., Wan L., Li G.X.,2006. Study on the depositional system of the Sanshui Basin, Guangdong, China. Geological Bulletin of China, 25(9), 1175-1183. https://doi.org/10.3969/j.issn.1671-2552.2006.09.032 (in Chinese with English abstract).
[21] Hou M.C., Chen H.D., Tian J.C., Wan L.,2007. Sedimentary facies and palaeogeography of the Sanshui Basin, Guangdong during the Paleogene. Sedimentary Geology and Tethyan Geology, 27(2), 37-44. https://doi.org/10.3969/j.issn.1009-3850.2007.02.006 (in Chinese with English abstract).
[22] Jackson F.D., Varricchio D.J.,2016. Fossil egg and eggshells from the Upper Cretaceous Hell Creek Formation, Montana. Journal of Vertebrate Paleontology, 36(5), e1185432. https://doi.org/10.1080/02724634.2016.1185432.
[23] Kohring R., Hirsch K.F.,1996. Crocodilian and avian eggshells from the middle Eocene of the Geiseltal, Eastern Germany. Journal of Vertebrate Paleontology, 16(1), 67-80. https://doi.org/10.1080/02724634.1996.10011285.
[24] Li C., Wu X.C., Rufolo S.J.,2019. A new crocodyloid (Eusuchia: Crocodylia) from the Upper Cretaceous of China. Cretaceous Research, 94, 25-39. https://doi.org/10.1016/j.cretres.2018.09.015.
[25] Liang H.Y., Sun W.D., Mo J.H., Hu G.Q., Zeng T., Allen C.M.,2008. Study on the structural-magmatic events and the formation of ore deposits in the Sanshui Basin, central Guangdong Province. Geology in China, 35(6), 1121-1128. https://doi.org/10.3969/j.issn.1000-3657.2008.06.009 (in Chinese with English abstract).
[26] López-Martínez N., Vicens E., 2012. A new peculiar dinosaur egg, Sankofa pyrenaica oogen. nov. oosp. nov. from the Upper Cretaceous coastal deposits of the Aren Formation, south-central Pyrenees, Lleida, Catalonia, Spain. Palaeontology, 55(2), 325-339. https://doi.org/10.1111/j.1475-4983.2011.01114.x.
[27] Lü J.C.,2003. A new oviraptorosaurid (Theropoda: Oviraptorosauria) from the Late Cretaceous of southern China. Journal of Vertebrate Paleontology, 22(4), 871-875. https://doi.org/10.1671/0272-4634(2002)022[0871:ANOTOF]2.0.CO;2.
[28] Lü J.C., Azuma Y., Huang D., Noda Y., Qiu L.C., 2006. New troodontid dinosaur egg from the Heyuan Basin of Guangdong Province, southern China. In: Lü, J.C., Kobayashi, Y., Huang, D., Lee, Y.N. (Eds.), Papers from the 2005 Heyuan International Dinosaur Symposium. Beijing: Geological Publishing House, pp. 11-18.
[29] Lü J.C., Chen R.J., Brusatte S.L., Zhu Y.X., Shen C.Z., 2016. A Late Cretaceous diversification of Asian oviraptorid dinosaurs: Evidence from a new species preserved in an unusual posture. Scientific Reports, 6, 35780. https://doi.org/10.1038/srep35780.
[30] Lü J.C., Li G.Q., Kundrát M., Lee Y.N., Sun Z.Y., Kobayashi Y., Shen C.Z., Teng F.F., Liu H.F., 2017. High diversity of the Ganzhou Oviraptorid Fauna increased by a new “cassowary-like” crested species. Scientific Reports, 7, 6393. https://doi.org/10.1038/s41598-017-05016-6.
[31] Lü J.C., Liu Y., Huang Z.Q., Yuan W.Q., Huang D.,2009. New discovery of Late Cretaceous tyrannosaurid tooth from the Heyuan Basin, Guangdong Province, southern China. Geological Bulletin of China, 28(6), 701-704. https://doi.org/10.3969/j.issn.1671-2552.2009.06.003 (in Chinese with English abstract).
[32] Lü J.C., Pu H.Y., Kobayashi Y., Xu L., Chang H.L., Shang Y.H., Liu D., Lee Y.N., Kundrát M., Shen C.Z., 2015. A new oviraptorid dinosaur (Dinosauria: Oviraptorosauria) from the Late Cretaceous of southern China and its paleobiogeographical implications. Scientific Reports, 5, 11490. https://doi.org/10.1038/srep11490.
[33] Lü J.C., Yi L.P., Zhong H., Wei X.F., 2013a. A new oviraptorosaur (Dinosauria: Oviraptorosauria) from the Late Cretaceous of southern China and its paleoecological implications. PLoS ONE, 8(11), e80557. https://doi.org/10.1371/journal.pone.0080557.
[34] Lü J.C., Yi L.P., Zhong H., Wei X.F., 2013b. A new somphospondylan sauropod (Dinosauria, Titanosauriformes) from the Late Cretaceous of Ganzhou, Jiangxi Province of southern China. Acta Geologica Sinica (English Edition), 87(3), 678-685. https://doi.org/10.1111/1755-6724.12079.
[35] Lü J.C., Zhang B.K.,2005. A new oviraptorid (Theropod: Oviraptorosauria) from the Upper Cretaceous of the Nanxiong Basin, Guangdong Province of southern China. Acta Palaeontologica Sinica, 44(3), 412-422. https://doi.org/10.3969/j.issn.0001-6616.2005.03.005 (in Chinese with English abstract).
[36] Mikhailov K.E.,1994. Theropod and protoceratopsian dinosaur eggs from the Cretaceous of Mongolia and Kazakhstan.Paleontological Journal, 28(2), 101-120.
[37] Moreno-Azanza,M., Canudo, J.I., Gasca, J.M., 2014. Unusual theropod eggshells from the Early Cretaceous Blesa Formation of the Iberian Range, Spain. Acta Palaeontologica Polonica, 59(4), 843-854. https://doi.org/10.4202/app.2012.0069.
[38] Norell M.A., Clark J.M., Demberelyin D., Rhinchen B., Chiappe L.M., Davidson A.R., McKenna M.C., Altangerel P., Novacek M.J., 1994. A theropod dinosaur embryo and the affinities of the Flaming Cliffs dinosaur eggs. Science, 266(5186), 779-782. https://doi.org/10.1126/science.266.5186.779.
[39] Oliveira C.E.M., Santucci R.M., Andrade M.B., Fulfaro V.J., Basílio J.A.F., Benton M.J., 2011. Crocodylomorph eggs and eggshells from the Adamantina Formation (Bauru Group), Upper Cretaceous of Brazil. Palaeontology, 54(2), 309-321. https://doi.org/10.1111/j.1475-4983.2010.01028.x.
[40] Panadès I.B.X., Patnaik R., 2009. A complete crocodylian egg from the upper Miocene (Chinji Beds) of Pakistan and its palaeobiographical implications.PalArchs Journal of Vertebrate Palaeontology 6(1): 1-8.
[41] Prasad G.V.R., Sharma, A., Verma, O., Khosla, A., Singh, L.R., Priyadarshini, R., 2015. Testudoid and crocodiloid eggshells from the Upper Cretaceous Deccan Intertrappean Beds of Central India. Comptes Rendus Palevol, 14(6), 513-526. https://doi.org/10.1016/j.crpv.2015.05.006.
[42] Qiu L.C., Huang D., 2001. Dinosaur fossils from the Heyuan Basin, Guangdong Province, China. In: Proceedings of the Eighth Annual Meeting of the Chinese Society of Vertebrate Paleontology. Beijing: China Ocean Press, pp. 59-63 (in Chinese with English abstract).
[43] Russo J., Mateus O., Marzola M., Balbino A., 2017. Two new ootaxa from the Late Jurassic: The oldest record of crocodylomorph eggs, from the Lourinhã Formation, Portugal. PLoS ONE, 12(3), e171919. https://doi.org/10.1371/journal.pone.0171919.
[44] Sun Yat-sen University., 1976. First discovery of dinosaur eggs from red deposits in Baihedong Group, Guangzhou.Acta Scientiarum Naturalium Universitatis Sunyatseni, 2, 137-138.
[45] Tanaka K., Lü J.C., Liu Y., Huang Z.Q., Kobayashi Y., Huang D., Zelenitsky D.K., 2012. Statistical approach for classification of dinosaur eggs from the Heyuan Basin at the northeast of Guangdong Province. Acta Geologica Sinica (English Edition), 86(2), 294-303. https://doi.org/10.1111/j.1755-6724.2012.00659.x.
[46] Tanaka K., Zelenitsky D.K., Therrien F., Ikeda T., Kubota K., Saegusa H., Tanaka T., Ikuno K.,2020. Exceptionally small theropod eggs from the Lower Cretaceous Ohyamashimo Formation of Tamba, Hyogo Prefecture, Japan. Cretaceous Research, 114, 104519. https://doi.org/10.1016/j.cretres.2020.104519.
[47] Tang Z.Y.,1994. Cretaceous-Eogene rift valley-type volcanism in Sanshui Basin, Guangdong.Guangdong Geology, 9(1), 49-57 (in Chinese).
[48] Wang D.Y., Zhou S.Q., 1995. The discovery of new typical dinosaur egg fossils in Xixia Basin.Henan Geology, 13(4), 262-267 (in Chinese with English abstract).
[49] Wang N.S., Kuang H.W., Liu Y.Q., Peng N., Xu H., Zhang P., Wang M.W., Wang B.H., An W.,2015. Taphonomic characteristics of the Late Cretaceous cluster dinosaur fossils in eastern China and comparison between domestic and abroad. Journal of Palaeogeography (Chinese Edition), 17(5), 593-610. https://doi.org/10.7605/gdlxb.2015.05.049 (in Chinese with English abstract).
[50] Wang Q., Li Y.G., Zhu X.F., Fang K.Y., Wang X.L., 2018. New ootype prismatoolithids from the Late Cretaceous, Laiyang Basin and its significance. Vertebrata PalAsiatica, 56(3), 264-272. https://doi.org/10.19615/j.cnki.1000-3118.180708.
[51] Wang Q., Wang X.L., Zhao Z.K., Jiang Y.G.,2010. A new oogenus of Elongatoolithidae from the Upper Cretaceous Chichengshan Formation of Tiantai Basin, Zhejiang Province. Vertebrata PalAsiatica, 48(2), 111-118. https://doi.org/10.3969/j.issn.1000-3118.2010.02.003 (in Chinese with English abstract).
[52] Wang Q., Zhao Z.K., Wang X.L., Li N., Zou S.L., 2013a. A new form of Elongatoolithidae, Undulatoolithus pengi oogen. et oosp. nov. from Pingxiang, Jiangxi, China. Zootaxa, 3746(1), 194-200. https://doi.org/10.11646/zootaxa.3746.1.9.
[53] Wang R., Jiang B.Y.,2021. Reconstruction of the Late Cretaceous paleoriver systems in China and their effects on distribution of hadrosaurs. Journal of Palaeogeography (Chinese Edition), 23(3), 581-599. https://doi.org/10.7605/gdlxb.2021.03.028 (in Chinese with English abstract).
[54] Wang S., Sun C.K., Sullivan C., Xu X., 2013b. A new oviraptorid (Dinosauria: Theropoda) from the Upper Cretaceous of southern China. Zootaxa, 3640(2), 242-257. https://doi.org/10.11646/zootaxa.3640.2.7.
[55] Wang X.F., Chen X.H., Zhang Z.L., Li X.B., 2004. Stratigraphic Division and Correlation of Each Geologic Period in China. Beijing: Geological Publishing House (in Chinese).
[56] Wang X.L., Wang Q., Jiang S.X., Cheng X., Zhang J.L., Zhao Z.K., Jiang Y.G., 2012. Dinosaur egg faunas of the Upper Cretaceous terrestrial red beds of China and their stratigraphical significance.Journal of Stratigraphy, 36(2), 400-416 (in Chinese with English abstract).
[57] Wei X.F., Pu H.Y., Xu L., Di Liu, Lü J.C., 2013. A new oviraptorid dinosaur (Theropoda: Oviraptorosauria) from the Late Cretaceous of Jiangxi Province, southern China. Acta Geologica Sinica (English Edition), 87(4), 899-904. https://doi.org/10.1111/1755-6724.12098.
[58] Wen C.H., Liu X.M., Lü B., Mao X.G., Chen J.S., Hou S.M., Zhou Z.B., Hou J.L., Wu H.B.,2016. The Cretaceous redbeds in Shicheng Basin, Jiangxi Province: Pedogenic and paleoenvironmental characteristics. Quaternary Sciences, 36(6), 1403-1416. https://doi.org/10.11928/j.issn.1001-7410.2016.06.06 (in Chinese with English abstract).
[59] Xing L.D., Harris J.D., Dong Z.M., Lin Y.L., Chen W., Guo S.B., Ji Q.,2009. Ornithopod (Dinosauria: Ornithischia) tracks from the Upper Cretaceous Zhutian Formation in the Nanxiong Basin, Guangdong, China and general observations on large Chinese ornithopod footprints. Geological Bulletin of China, 28(7), 829-843. https://doi.org/10.3969/j.issn.1671-2552.2009.07.003 (in Chinese with English abstract).
[60] Xing L.D., Lockley M.G., Li D.L., Klein H., Ye Y.,Scott Persons IV, W., Ran, H., 2017. Late Cretaceous ornithopod-dominated, theropod, and pterosaur track assemblages from the Nanxiong Basin, China: New discoveries, ichnotaxonomy, and paleoecology. Palaeogeography, Palaeoclimatology, Palaeoecology, 466, 303-313. https://doi.org/10.1016/j.palaeo.2016.11.035.
[61] Xing L.D., Niu K.C., Lockley M.G., Klein H., Romilio A.,Scott Persons IV, W., Brusatte, S.L., 2019. A probable tyrannosaurid track from the Upper Cretaceous of southern China. Science Bulletin, 64(16), 1136-1139. https://doi.org/10.1016/j.scib.2019.06.013.
[62] Xing L.D., Niu K.C., Ma W.S., Zelenitsky D.K., Yang T.R., Brusatte S.L.,2022b. An exquisitely preserved in-ovo theropod dinosaur embryo sheds light on avian-like prehatching postures. iScience, 25(1), 103516. https://doi.org/10.1016/j.isci.2021.103516.
[63] Xing L.D., Niu K.C., Wang D.H., Marquez A.P.,2021. A partial articulated hadrosaurid skeleton from the Maastrichtian (Upper Cretaceous) of the Ganzhou area, Jiangxi Province, China. Historical Biology, 33(10), 2256-2259. https://doi.org/10.1080/08912963.2020.1782397.
[64] Xing L.D., Niu K.C., Yang T.R., Wang D.H., Miyashita T., Mallon J.C., 2022a. Hadrosauroid eggs and embryos from the Upper Cretaceous (Maastrichtian) of Jiangxi Province, China. BMC Ecology and Evolution, 22(1), 60. https://doi.org/10.1186/s12862-022-02012-x.
[65] Xu L., Xie J.F., Zhang S.K., Choi S., Kim N.H., Gao D.S., Jin X.S., Jia S.H., Gao Y.L.,2022. Fossil turtle eggs from the Upper Cretaceous Gaogou Formation, Xiaguan-Gaoqiu Basin, Neixiang County, Henan Province, China: Interpretation of the transformation from aragonite to calcite in fossil turtle eggshell. Cretaceous Research, 134, 105166. https://doi.org/10.1016/j.cretres.2022.105166.
[66] Xu X., Han F.L.,2010. A new oviraptorid dinosaur (Theropoda: Oviraptorosauria) from the Upper Cretaceous of China. Vertebrata PalAsiatica, 48(1), 11-18. https://doi.org/10.3969/j.issn.1000-3118.2010.01.002.
[67] Young C.C.,1954. Fossil reptilian eggs from Laiyang, Shantung, China.Acta Scientia Sinica, 3(4), 505-522.
[68] Young C.C.,1965a. Fossil eggs from Nanhsiung, Kwangtung and Kanchou, Kiangsi.Vertebrata PalAsiatica, 9(2), 141-159 (in Chinese), 159-189 (in English).
[69] Young C.C.,1965b. Note on the reptilian remains from Nanhsiung, Kwangtung.Vertebrata PalAsiatica, 9(3), 292-297.
[70] Young C.C., Chow M.Z., 1962. Some reptilian fossils from the “red beds” of Kwangtung and Chekiang.Vertebrata PalAsiatica, 6(2), 130-137.
[71] Yu C.T., Fan X.J.,2022. Distribution of dinosaur egg fossils and occurrence stratigraphic characteristics of Meilin-Maodian, Ganzhou Basin. Journal of Hebei GEO University, 45(4), 6-12. https://doi.org/10.13937/j.cnki.hbdzdxxb.2022.04.002 (in Chinese with English abstract).
[72] Yu J.X., Sun M.R., Sun S.Y., Mao S.Z., 1981. Dinoflagellates and acritarchs from Dalangshan Formation and their significance in explaining the depositional environment.Oil and Gas Geology, 2(3), 254-264 (in Chinese with English abstract).
[73] Yu X.Q.,1998. Characteristics of dinosaur fossils from southern Anhui and their significance for stratigraphic division.Regional Geology of China, 17(3), 278-284 (in Chinese with English abstract).
[74] Yuan Y.R., Wang B.G., Tang X.Z., Lai X.J., Hou H.M., Fan S.Q., Liu Z.Y., 1994. Cretaceous-Early Tertiary paleolatitude migration in the Sanshui Basin of South China and the evolution of the South China Sea. Tectonophysics, 235(1-2), 117-129. https://doi.org/10.1016/0040-1951(94)90020-5.
[75] Zelenitsky D.K., Hills L.V., 1996. An egg clutch of Prismatoolithus levis oosp. nov. from the Oldman Formation (Upper Cretaceous), Devil's Coulee, southern Alberta. Canadian Journal of Earth Sciences, 33(8), 1127-1131. https://doi.org/10.1139/e96-085.
[76] Zeng D.M., Zhang J.J., 1979. On the dinosaurian eggs from the western Dongting Basin, Hunan.Vertebrata PalAsiatica, 17(2), 131-136 (in Chinese with English abstract).
[77] Zhang W., Huang P.Y.,1999. The strata with fossil dinosaur eggs and their distribution in Guangdong. Journal of Stratigraphy, 23(2), 89-95. https://doi.org/10.3969/j.issn.0253-4959.1999.02.002 (in Chinese with English abstract).
[78] Zhang X.Q.,1984. On the subdivision and correlation of the Cretaceous System of Sanshui Basin of Guangdong. Petroleum Geology and Experiment, 6(1), 32-40. https://doi.org/10.11781/sysydz198401032 (in Chinese with English abstract).
[79] Zhang X.Q., Li G., 2015. Discussion on geological age of the Pingling Member of Shanghu Formation in the Nanxiong Basin, Guangdong Province.Journal of Stratigraphy, 39(1), 74-80 (in Chinese with English abstract).
[80] Zhang X.Q., Li G., Yang R.L., Li H.M.,2008. Late Cretaceous ostracods from the Sanshui Basin of Guangdong. Acta Micropalaeontologica Sinica, 25(2), 132-165. https://doi.org/10.3969/j.issn.1000-0674.2008.02.004 (in Chinese with English abstract).
[81] Zhang X.Q., Ling Q.X., Lin J.N., 2005. Research status quo about the red beds of the Heyuan Basin, Guangdong.Journal of Stratigraphy, 29(S1), 602-607 (in Chinese with English abstract).
[82] Zhang X.Q., Zhou X.P., Chen X.Y., 1993. Atlas of Cretaceous-Tertiary Drilling Strata Division and Correlation in Sanshui Basin. Beijing: Ocean Publishing House, pp. 1-136.
[83] Zhao H., Zhao Z.K., 1999. A new form of elongatoolithid dinosaur eggs from the Lower Cretaceous Shahai Formation of Heishan, Liaoning Province.Vertebrata PalAsiatica, 37(4), 278-284.
[84] Zhao Z.K.,1975. Microscopic structure of dinosaur egg fossils in Nanxiong, Guangdong Province (I) — also on the classification of dinosaur egg fossils.Vertebrata PalAsiatica, 13(2), 105-117 (in Chinese).
[85] Zhao Z.K.,2000. Nesting behavior of dinosaurs as interpreted from the Chinese Cretaceous dinosaur eggs.Paleontological Society of Korea, Special Publication, 4, 115-126.
[86] Zhao Z.K., Li R., 1993. First record of Late Cretaceous hypsilophodontid eggs from Bayan Manduhu, Inner Mongolia.Vertebrata PalAsiatica, 31(2), 77-84.
[87] Zhao Z.K., Wang Q., Zhang S.K., 2015. Dinosaur Eggs. Vertebrate Paleontology of China, Volume 7. Beijing: Science Press, pp. 1-168. (in Chinese).
[88] Zhao Z.K., Ye J., Li H.M., Zhao Z.H., Yan Z., 1991. Extinction of the dinosaurs across the Cretaceous-Tertiary boundary in Nanxiong Basin, Guangdong Province.Vertebrata PalAsiatica, 29(1), 1-12.
[89] Zhu X.F., Fang K.Y., Wang Q., Lu X.H., Wu W.Q., Du Y.L., Huang Z.Q., Wang X.L., 2019. The first Stalicoolithus shifengensis discovered in a clutch from the Sanshui Basin, Guangdong Province. Vertebrata PalAsiatica, 57(1), 77-83. https://doi.org/10.19615/j.cnki.1000-3118.180110.