甘肃永昌乌拉尔世<italic>Sphenopteris</italic>(楔羊齿)新记录: 兼论晚古生代<italic>Sphenopteris</italic>多样性及古地理演化<sup>*</sup>

doi:10.7605/gdlxb.2023.02.029

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Abstract Although Sphenopteris species were widespread in the Late Paleozoic around the world,the systematic position and evolutionary characteristics of this genus still need to be addressed,and microstructures such as epidermis and stomata have only been demonstrated in very limited species. We describe two well-preserved fossil species with microstructures from the Cisuralian Shanxi Formation(Early Permian)of Yongchang,Gansu,China,including Sphenopteris yongchangensis sp. nov. and Sphenopteris nystroemii. Pinnules of S. yongchangensis sp. nov. were fan-shaped,with their base connected,lateral veins bifurcate once,and elliptical stomata are amphistomatic. Pinnules of S. nystroemii were fusiform with round top,lateral veins bifurcate once and reach leaf margin,and its stomata were distributed in interveinal area. It is more reasonable to assign the present two species of Sphenopteris to the order Lyginopteridales by detailed comparative analyses. The diversity and palaeogeographic evolution of Sphenopteris species are then considered,based on 269 occurrence records of the genus from the Late Paleozoic,in combination with morphological and anatomical characteristics and biogeographic distribution pattern. The results show that the records of Sphenopteris were widely distributed in the middle-low latitudes worldwide from the Pennsylvania to Cisuralian,indicating high variation and differentiation of Sphenopteris-type leaves. It is speculated that Sphenopteris species migrated from South China to North China during Late Devonian to Permian,and the diversity pattern was not only closely related to humid climate,but also coupled with the palaeogeographic characteristics of this genus.

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	Articles by authors
	WANG Xuelian
	XIONG Conghui
	SUN Bainian

Key words： Late Paleozoic Sphenopteris palaeogeographic distribution diversity

Received: 01 December 2022

ZTFLH:

Q914.7

Fund:National Natural Science Foundation of China(Nos.42202029,41802008,41888101),Second Tibetan Plateau Scientific Expedition and Research Program(No.2019QZKK0707),and the Natural Science Foundation of Gansu Province(Nos. 21JR7RA061,20JR10RA646)

Corresponding Authors: XIONG Conghui,born in 1986,is a senior engineer at School of Earth Sciences,Lanzhou University. Her research focuses on the diversity of Paleozoic vascular land plants. E-mail: xiongch@lzu.edu.cn.

About author: WANG Xuelian,born in 1991,is an assistant researcher of paleobotany and paleoenvironments,at Northwest Institute of Eco?Environment and Resources,Chinese Academy of Sciences. Her research focuses on paleobotany and climate changes. E-mail: wangxuelian@nieer.ac.cn.

Cite this article:

WANG Xuelian,XIONG Conghui,SUN Bainian. New records of Sphenopteris from the Cisuralian of Yongchang,Gansu Province,with a discussion on diversity and palaeogeographic evolution of this genus during the Late Paleozoic[J]. JOPC, 2023, 25(2): 471-484.

URL:

http://journal09.magtechjournal.com/gdlxb/EN/10.7605/gdlxb.2023.02.029 OR http://journal09.magtechjournal.com/gdlxb/EN/Y2023/V25/I2/471

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